Based on the review of micromammals associations (Soricomorphs, Erinaceomorphs, Chiropters and Rodents) from several sites in southern France (Arago Cave, Lazaret Cave, Moula-Guercy cave) and from the study of an original deposit in south-west of the Iberian Peninsula (Maltravieso cave), the main aim of this work is to reconstruct the evolution of palaeoenvironments and palaeoclimates in southern Europe, during Middle Pleistocene between 600 and 130 ka BP. In order to assess the reliability of palaeoecological reconstructions, taphonomic analysis have been performed on micromammals bones, and have allowed to highlight the involvement of opportunistic and specialized owls in the origin of accumulations. Palaeoecological analysis, based on the use of different types of methods, showed that the alternating glacial cycles, characterized by cold or cool climates and open spaces, and interglacial, more temperate with closed environments, had an essential impact on the distribution of micromammals species in southern Europe, in relation to geography and topography of each site. "Non-analogue" communities are identified, combining temperate and cold zones taxa, reflecting the role of refugia area of these regions. They also show that dispersal events have affected palaeobiodiversity during the Middle Pleistocene, particularly in the areas of maximum expansion of species during the glacial phases.
